Cynoglossum

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Federico Selvi - One of the best experts on this subject based on the ideXlab platform.

  • multiple origins for hound s tongues Cynoglossum l and navel seeds omphalodes mill the phylogeny of the borage family boraginaceae s str
    2013
    Co-Authors: Maximilian Weigend, Federico Selvi, Federico Luebert, Grischa Brokamp, Hartmut H. Hilger
    Abstract:

    Abstract Recent studies all indicated that both the affinities and subdivision of Boraginaceae s.str. are unsatisfactorily resolved. Major open issues are the placement and affinities of Boraginaceae s.str. in Boraginales and the major clades of the family, with especially the large tribes Cynoglosseae and Eritrichieae repeatedly retrieved as non-monophyletic groups, and the doubtful monophyly of several larger genera, especially Cynoglossum and Omphalodes . The present study addresses and solves these questions using two plastid markers ( trnL – trnF , rps 16) on the basis of a sampling including 16 outgroup taxa and 172 ingroup species from 65 genera. The phylogeny shows high statistical support for most nodes on the backbone and on the individual clades. Boraginaceae s.str. are sister to African Wellstediaceae, Wellstediaceae–Boraginaceae s.str. is sister to African Codonaceae. Echiochileae are retrieved as sister to the remainder of Boraginaceae s.str., which, in turn, fall into two major clades, the Boragineae–Lithospermeae (in a well-supported sister relationship) and the Cynoglosseae s.l. (including Eritrichieae). Cynoglosseae s.l. is highly resolved, with Trichodesmeae (incl. Microcaryum , Lasiocaryum ) as sister to the remainder of the group. Eritrichieae s.str. ( Eritrichium , Hackelia , Lappula ) are resolved on a poorly supported polytomy together with the Omphalodes -clade (incl. Myosotidium , Cynoglossum p.p.), and the Mertensia -clade (incl. O. scorpioides , Asperugo ). The Myosotideae ( Myosotis , Trigonotis , Pseudomertensia ) are retrieved in a well-supported sister-relationship to the core-Cynoglosseae, the latter comprising all other genera sampled. Cynoglossum is retrieved as highly para- and polyphyletic, with a large range of generic segregates embedded in Cynoglossum , but other species of Cynoglossum are sister to Microula or to the American “Eritrichieae” ( Cryptantha and allied genera). Representatives of the genus Cynoglossum in its current definition are segregated onto six independent lineages, members of Omphalodes onto three independent lineages. At least 11 of the genera here sampled are deeply nested in other genera. The data show that individual details of nutlet morphology (e.g., winged margins, glochidia) are highly homoplasious. Conversely, a complex of nutlet characters (e.g., characters of the gynobase and cicatrix together with nutlet orientation and sculpturing) tends to circumscribe natural units. Geographical distribution of major clades suggests that the family originated in Africa and western Asia and radiated to eastern Eurasia, with several independent dispersal events into Australia and the New World.

  • a synopsis of the genus Cynoglossum boraginaceae cynoglosseae in italy
    2012
    Co-Authors: Federico Selvi, Karel Sutorý
    Abstract:

    Abstract Based on field and herbarium investigations, a systematic synthesis of the critical genus Cynoglossum (Boraginaceae) in Italy is provided. Types, diagnostic characters and original iconographies are given for each native species, together with a revised analytical key. Regional distributions are outlined and a list of selected vouchers is included. Nine species are native to the Italian territory: Cynoglossum barbaricinum, Cynoglossum cheirifolium, Cynoglossum clandestinum, C. columnae, C. creticum, C. magellense, C. montanum, C. nebrodense and C. officinale. A new subspecies, named C. nebrodense ssp. lucanum, is described from some calcareous massifs in S Basilicata and N Calabria. This taxon is morphologically and geographically distinct from the typical C. nebrodense s.str. In addition, C. dioscoridis and C. germanicum, often reported from Italy, cannot be confirmed and are apparently to be excluded from the Italian flora. Old herbarium records of C. amabile Stapf & Drummond and C. coelestinum...

  • typification of names of euro mediterranean taxa of boraginaceae described by italian botanists
    2009
    Co-Authors: Federico Selvi, Lorenzo Cecchi
    Abstract:

    Nomenclatural types for 26 names in the Boraginaceae genera Caccinia Savi, Cerinthe L., Cynoglossum L., Echium L., Lithospermum L., Myosotis L., Omphalodes Mill. and Onosma L., validly published by Italian botanists from Domenico Viviani to Adriano Fiori, are newly designated. Most of these names apply to taxa of the Eurasiatic flora which are accepted in the modern taxonomic and floristic literature. The new combinations Buglossoides incrassata (Guss.) I.M. Johnst. subsp. splitgerberi (Guss.) E. Zippel & Selvi and Omphalodes rupestris Rupr. ex Boiss. subsp. lojkae (Sommier & Levier) Selvi are proposed, and the new species Myosotis graui Selvi, previously known under the misapplied name M. ambigens (Beg.) Grau, is described.

  • epitypification and identity of Cynoglossum montanum l boraginaceae
    2008
    Co-Authors: Federico Selvi
    Abstract:

    An epitype for Cynoglossum montanum, a Linnaean name applied to a controversial species described from Italy, is designated in view of the poor characterization of its lectotype, an illustration published in the year 1616 based on plants from the central Apennines. The morphological analysis of the epitype and other herbarium material from the Italian peninsula allowed a better definition of diagnostic characters with respect to the closely related and broadly sympatric C. officinale, to which is connected by intermediate forms.

Stier Victoria - One of the best experts on this subject based on the ideXlab platform.

  • Supplementary material 1 from: Hilger H, Greuter W, Stier V (2015) Taxa and names in Cynoglossum sensu lato (Boraginaceae, Cynoglosseae): an annotated, synonymic inventory, with links to the protologues and mention of original material. Biodiversity Data Journal 3: e4831. https://doi.org/10.3897/BDJ.3.e4831
    2015
    Co-Authors: Hilger Hartmut, Greuter Werner, Stier Victoria
    Abstract:

    Nomenclatural database for Cynoglossum sensu lato - a structured, fully searchable Hypertext Markup Language (HTML) file with names connected by internal links and, where appropriate, linked to external sources

  • Taxa and names in Cynoglossum sensu lato (Boraginaceae, Cynoglosseae): an annotated, synonymic inventory, with links to the protologues and mention of original material
    2015
    Co-Authors: Hilger Hartmut, Greuter Werner, Stier Victoria
    Abstract:

    An inventory is presented of all names so far validly published in Cynoglossum sensu lato and its segregate genera: Adelocaryum, Afrotysonia, Kuschakewiczia, Lindelofia, Mattiastrum, Paracaryum, Rindera, Solenanthus, Trachelanthus, and their synonyms. Names and designations that were not validly published in the cited place, and later isonyms, are accounted for when they have been included in the International Plant Name Index (IPNI). Problems with IPNI entries, including errors and omissions, are discussed, and the hope is expressed that the present inventory may be of use for fixing them. The inventory, generated from a list of structured data, is presented in two Supplements, as a searchable HTML document comprising a sequence of entries with internal cross-links and links to external sources, in particular to protologues accessible online or, copyright restrictions permitting, made available as scanned documents via DOIs, and as machine-readible file. With minor exceptions, all names have been verified in their original place of publication, and all were nomenclaturally assessed. Colour coding is used to distinguish between names (in green) pertaining to Cynoglossum sensu lato, for which complete synonymies are provided; and names (in orange) pertaining to other genera but published under Cynoglossum or its segregates. They are listed together with their basionym and the corresponding correct name (if it exists), but without complete synonymy. Acceptable, potentially correct names appear in bold-face type, both under a broadly defined Cynoglossum (for which purpose validation of 81 new combinations and the name of 1 new species was necessary) and under one or more of its segregates. When a name was published for a new taxon, original material is indicated, usually by direct quotation from the protologue. New type designations are exceptional (two cases), whereas former type designations are cited whenever known. Furthermore, types and original specimens, especially when their digital images are available online, are mentioned with their locations and accession numbers. Comments are added whenever appropriate, especially to explain nomenclatural assessments that are not self-evident

  • an annotated, synonymic inventory, with links to the protologues and mention of original material
    2015
    Co-Authors: Hilger, Hartmut H., Greuter Werner, Stier Victoria
    Abstract:

    Background An inventory is presented of all names so far validly published in Cynoglossum sensu lato and its segregate genera: Adelocaryum, Afrotysonia, Kuschakewiczia, Lindelofia, Mattiastrum, Paracaryum, Rindera, Solenanthus, Trachelanthus, and their synonyms. Names and designations that were not validly published in the cited place, and later isonyms, are accounted for when they have been included in the International Plant Name Index (IPNI). Problems with IPNI entries, including errors and omissions, are discussed, and the hope is expressed that the present inventory may be of use for fixing them. New information The inventory, generated from a list of structured data, is presented in two Supplements, as a searchable HTML document comprising a sequence of entries with internal cross-links and links to external sources, in particular to protologues accessible online or, copyright restrictions permitting, made available as scanned documents via DOIs, and as machine- readible file. With minor exceptions, all names have been verified in their original place of publication, and all were nomenclaturally assessed. Colour coding is used to distinguish between names (in green) pertaining to Cynoglossum sensu lato, for which complete synonymies are provided; and names (in orange) pertaining to other genera but published under Cynoglossum or its segregates. They are listed together with their basionym and the corresponding correct name (if it exists), but without complete synonymy. Acceptable, potentially correct names appear in bold-face type, both under a broadly defined Cynoglossum (for which purpose validation of 81 new combinations and the name of 1 new species was necessary) and under one or more of its segregates. When a name was published for a new taxon, original material is indicated, usually by direct quotation from the protologue. New type designations are exceptional (two cases), whereas former type designations are cited whenever known. Furthermore, types and original specimens, especially when their digital images are available online, are mentioned with their locations and accession numbers. Comments are added whenever appropriate, especially to explain nomenclatural assessments that are not self-evident

Hilger Hartmut - One of the best experts on this subject based on the ideXlab platform.

Tom J De Jong - One of the best experts on this subject based on the ideXlab platform.

  • Bidirectional selection on threshold size for flowering in Cynoglossum officinale (hound's-tongue)
    1995
    Co-Authors: Renate A. Wesselingh, Tom J De Jong
    Abstract:

    Previous studies have demonstrated that many facultative biennials show variation in threshold size for flowering. In order to quantify the genetic variation for this character, we performed a two-way artificial selection experiment on threshold size in Cynoglossum officinale. The parental generation, established from seed from a natural population, showed large variation in threshold sizes (2.6- 13.4 g). After one generation of selection for low threshold sizes all plants in the F1 flowered at sizes below 3.2 g. In the high selection line none of the F1 plants under 3.6 g flowered, and there were very large nonflowering plants (up to 17.2 g). By interpreting the logistic regression of flowering probability on plant size as a cumulative frequency distribution of threshold sizes, we could derive the frequency distributions of threshold sizes in each generation. These were used to estimate the narrow-sense heritabilities by the standard procedure for artificial selection on a quantitative character. This gave 'heritabilities' of 0.35 and 0.32 for the high and low selection lines, respectively. A previously constructed optimization model, applied to the population in our study area, predicted strong selection against low threshold sizes, but a fairly equal fitness for threshold sizes around and above the optimum. This flat fitness profile may explain why the natural population of Cynoglossum officinale harbours extensive genetic variation for this character.

  • Geographical variation in threshold size for flowering in Cynoglossum officinale
    1993
    Co-Authors: Renate A. Wesselingh, Tom J De Jong, Peter G. L. Klinkhamer, Marinke J. Van Dijk, Els G. M. Schlatmann
    Abstract:

    We investigated variation in two traits that determine generation time, cold- and size-requirement for flowering, within and among EuroPean populations for the monocarpic perennial Cynoglossum officinale. When grown in an experimental garden in Leiden, no annual individuals were found among plants originating from 22 locations; all plants were biennial under nutrient-rich growing conditions. In a controlled-environment experiment, in which plants received an artificial cold treatment, flowering probability increased gradually with plant size for plants from two natural populations, signifying a large within-population variation in threshold size for flowering. The relationships between plant size and flowering of these two groups were significantly different: plants from Holkham (England) had much higher threshold sizes than plants from Meijendel (The Netherlands). Three plant groups originating from botanic gardens showed a sharp increase in flowering probability with size, indicating less variation in threshold size. Significant differences existed among all five groups. Results indicate the possibility that natural selection acts upon threshold size for flowering in Cynoglossum officinale

Hartmut H. Hilger - One of the best experts on this subject based on the ideXlab platform.

  • a revision of the genus Cynoglossum l boraginaceae juss in nepal and notes on the widespread asian species
    2015
    Co-Authors: Julia Konig, Hartmut H. Hilger, Mohan Siwakoti, Maximilian Weigend
    Abstract:

    Cynoglossum is one of the most complex genera of Boraginaceae, with both species limits and the delimitation of the genus and its generic segregates highly problematic. Several hundred taxa and more than one thousand names are currently assigned to the genus and its segregates. The geographical centres of distribution are found in Asia and the Mediterranean. The present study investigates the genus Cynoglossum for Nepal, based mainly on herbarium collections from Nepal and across Asia, including most types, and some living accessions in Bonn Botanical Gardens. Based on this revision a total of five species of Cynoglossum are recognised for Nepal: Cynoglossum amabile, C. furcatum, C. lanceolatum, C. microglochin and C. wallichii. Cynoglossum microglochin , a large-fruited taxon, is here for the first time reported from Nepal and is the only species restricted to the Himalayas. All other species are widespread. The widely used name Cynoglossum zeylanicum is not available and the specimens so identified belong to C. furcatum , which is shown to extend to Georgia (Caucasus) in the West. Species described under ParaCynoglossum and Cynoglossum, respectively, are found to be synonymous with each other, arguing for abandoning the problematic recognition of ParaCynoglossum as a distinct genus. Diagnostic characters are described, micromorphological characters are illustrated and a key for identifying the five species is provided.

  • taxa and names in Cynoglossum sensu lato boraginaceae cynoglosseae an annotated synonymic inventory with links to the protologues and mention of original material
    2015
    Co-Authors: Hartmut H. Hilger, Werner Greuter, Victoria Stier
    Abstract:

    Background An inventory is presented of all names so far validly published in Cynoglossum sensu lato and its segregate genera: Adelocaryum, Afrotysonia, Kuschakewiczia, Lindelofia, Mattiastrum, Paracaryum, Rindera, Solenanthus, Trachelanthus, and their synonyms. Names and designations that were not validly published in the cited place, and later isonyms, are accounted for when they have been included in the International Plant Name Index (IPNI). Problems with IPNI entries, including errors and omissions, are discussed, and the hope is expressed that the present inventory may be of use for fixing them.

  • multiple origins for hound s tongues Cynoglossum l and navel seeds omphalodes mill the phylogeny of the borage family boraginaceae s str
    2013
    Co-Authors: Maximilian Weigend, Federico Selvi, Federico Luebert, Grischa Brokamp, Hartmut H. Hilger
    Abstract:

    Abstract Recent studies all indicated that both the affinities and subdivision of Boraginaceae s.str. are unsatisfactorily resolved. Major open issues are the placement and affinities of Boraginaceae s.str. in Boraginales and the major clades of the family, with especially the large tribes Cynoglosseae and Eritrichieae repeatedly retrieved as non-monophyletic groups, and the doubtful monophyly of several larger genera, especially Cynoglossum and Omphalodes . The present study addresses and solves these questions using two plastid markers ( trnL – trnF , rps 16) on the basis of a sampling including 16 outgroup taxa and 172 ingroup species from 65 genera. The phylogeny shows high statistical support for most nodes on the backbone and on the individual clades. Boraginaceae s.str. are sister to African Wellstediaceae, Wellstediaceae–Boraginaceae s.str. is sister to African Codonaceae. Echiochileae are retrieved as sister to the remainder of Boraginaceae s.str., which, in turn, fall into two major clades, the Boragineae–Lithospermeae (in a well-supported sister relationship) and the Cynoglosseae s.l. (including Eritrichieae). Cynoglosseae s.l. is highly resolved, with Trichodesmeae (incl. Microcaryum , Lasiocaryum ) as sister to the remainder of the group. Eritrichieae s.str. ( Eritrichium , Hackelia , Lappula ) are resolved on a poorly supported polytomy together with the Omphalodes -clade (incl. Myosotidium , Cynoglossum p.p.), and the Mertensia -clade (incl. O. scorpioides , Asperugo ). The Myosotideae ( Myosotis , Trigonotis , Pseudomertensia ) are retrieved in a well-supported sister-relationship to the core-Cynoglosseae, the latter comprising all other genera sampled. Cynoglossum is retrieved as highly para- and polyphyletic, with a large range of generic segregates embedded in Cynoglossum , but other species of Cynoglossum are sister to Microula or to the American “Eritrichieae” ( Cryptantha and allied genera). Representatives of the genus Cynoglossum in its current definition are segregated onto six independent lineages, members of Omphalodes onto three independent lineages. At least 11 of the genera here sampled are deeply nested in other genera. The data show that individual details of nutlet morphology (e.g., winged margins, glochidia) are highly homoplasious. Conversely, a complex of nutlet characters (e.g., characters of the gynobase and cicatrix together with nutlet orientation and sculpturing) tends to circumscribe natural units. Geographical distribution of major clades suggests that the family originated in Africa and western Asia and radiated to eastern Eurasia, with several independent dispersal events into Australia and the New World.