The Experts below are selected from a list of 336 Experts worldwide ranked by ideXlab platform

Alain Krief - One of the best experts on this subject based on the ideXlab platform.

Adrian Kremer - One of the best experts on this subject based on the ideXlab platform.

Stephane Jeanmart - One of the best experts on this subject based on the ideXlab platform.

Eran Pichersky - One of the best experts on this subject based on the ideXlab platform.

  • pyrethric Acid of natural pyrethrin insecticide complete pathway elucidation and reconstitution in nicotiana benthamiana
    New Phytologist, 2019
    Co-Authors: Anthony L Schilmiller, Maarten A Jongsma, Henriette D L M Van Eekelen, Ric C H De Vos, Eran Pichersky
    Abstract:

    In the natural pesticides known as pyrethrins, which are esters produced in flowers of Tanacetum cinerariifolium (Asteraceae), the monoterpenoid acyl moiety is pyrethric Acid or Chrysanthemic Acid. We show here that pyrethric Acid is produced from chrysanthemol in six steps catalyzed by four enzymes, the first five steps occurring in the trichomes covering the ovaries and the last one occurring inside the ovary tissues. Three steps involve the successive oxidation of carbon 10 (C10) to a carboxylic group by TcCHH, a cytochrome P450 oxidoreductase. Two other steps involve the successive oxidation of the hydroxylated carbon 1 to give a carboxylic group by TcADH2 and TcALDH1, the same enzymes that catalyze these reactions in the formation of Chrysanthemic Acid. The ultimate result of the actions of these three enzymes is the formation of 10‐carboxyChrysanthemic Acid in the trichomes. Finally, the carboxyl group at C10 is methylated by TcCCMT, a member of the SABATH methyltransferase family, to give pyrethric Acid. This reaction occurs mostly in the ovaries. Expression in N. benthamiana plants of all four genes encoding aforementioned enzymes, together with TcCDS, a gene that encodes an enzyme that catalyzes the formation of chrysanthemol, led to the production of pyrethric Acid.

  • production of trans Chrysanthemic Acid the monoterpene Acid moiety of natural pyrethrin insecticides in tomato fruit
    Metabolic Engineering, 2018
    Co-Authors: Daniel B Lybrand, Stefan Bennewitz, Alain Tissier, Eran Pichersky
    Abstract:

    The pyrethrum plant, Tanacetum cinerariifolium (Asteraceae) synthesizes a class of compounds called pyrethrins that have strong insecticidal properties but are safe to humans. Class I pyrethrins are esters of the monoterpenoid trans-Chrysanthemic Acid with one of three jasmonic-Acid derived alcohols. We reconstructed the trans-Chrysanthemic Acid biosynthetic pathway in tomato fruits, which naturally produce high levels of the tetraterpene pigment lycopene, an isoprenoid which shares a common precursor, dimethylallyl diphosphate (DMAPP), with trans-Chrysanthemic Acid. trans-Chrysanthemic Acid biosynthesis in tomato fruit was achieved by expressing the chrysanthemyl diphosphate synthase gene from T. cinerariifolium, encoding the enzyme that uses DMAPP to make trans-chrysanthemol, under the control of the fruit specific promoter PG, as well as an alcohol dehydrogenease (ADH) gene and aldehyde dehydrogenase (ALDH) gene from a wild tomato species, also under the control of the PG promoter. Tomato fruits expressing all three genes had a concentration of trans-Chrysanthemic Acid that was about 1.7-fold higher (by weight) than the levels of lycopene present in non-transgenic fruit, while the level of lycopene in the transgenic plants was reduced by 68%. Ninety seven percent of the diverted DMAPP was converted to trans-Chrysanthemic Acid, but 62% of this Acid was further glycosylated. We conclude that the tomato fruit is an alternative platform for the biosynthesis of trans-Chrysanthemic Acid by metabolic engineering.

  • coexpression analysis identifies two oxidoreductases involved in the biosynthesis of the monoterpene Acid moiety of natural pyrethrin insecticides in tanacetum cinerariifolium
    Plant Physiology, 2018
    Co-Authors: Gaurav D Moghe, Krystle Wiegertrininger, Anthony L Schilmiller, Cornelius S Barry, Eran Pichersky
    Abstract:

    Flowers of Tanacetum cinerariifolium produce a set of compounds known collectively as pyrethrins, which are commercially important pesticides that are strongly toxic to flying insects but not to most vertebrates. A pyrethrin molecule is an ester consisting of either trans-Chrysanthemic Acid or its modified form, pyrethric Acid, and one of three alcohols, jasmolone, pyrethrolone, and cinerolone, that appear to be derived from jasmonic Acid. Chrysanthemyl diphosphate synthase (CDS), the first enzyme involved in the synthesis of trans-Chrysanthemic Acid, was characterized previously and its gene isolated. TcCDS produces free trans-chrysanthemol in addition to trans-chrysanthemyl diphosphate, but the enzymes responsible for the conversion of trans-chrysanthemol to the corresponding aldehyde and then to the Acid have not been reported. We used an RNA sequencing-based approach and coexpression correlation analysis to identify several candidate genes encoding putative trans-chrysanthemol and trans-chrysanthemal dehydrogenases. We functionally characterized the proteins encoded by these genes using a combination of in vitro biochemical assays and heterologous expression in planta to demonstrate that TcADH2 encodes an enzyme that oxidizes trans-chrysanthemol to trans-chrysanthemal, while TcALDH1 encodes an enzyme that oxidizes trans-chrysanthemal into trans-Chrysanthemic Acid. Transient coexpression of TcADH2 and TcALDH1 together with TcCDS in Nicotiana benthamiana leaves results in the production of trans-Chrysanthemic Acid as well as several other side products. The majority (58%) of trans-Chrysanthemic Acid was glycosylated or otherwise modified. Overall, these data identify key steps in the biosynthesis of pyrethrins and demonstrate the feasibility of metabolic engineering to produce components of these defense compounds in a heterologous host.

Paolo Righi - One of the best experts on this subject based on the ideXlab platform.