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C. Thomas Philbrick - One of the best experts on this subject based on the ideXlab platform.
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Contributions to the taxonomy of Rhyncholacis (Podostemaceae): Evidence of monophyly, description of a new species, and transfer of the monotypic Macarenia
Phytotaxa, 2018Co-Authors: C. Thomas Philbrick, Brad R. Ruhfel, Claudia Petean BoveAbstract:We conducted a phylogenetic study of neotropical subfamily Podostemoideae with a focus on Rhyncholacis and the monotypic Macarenia using molecular data (plastid: rbcL , trnL intron; nuclear: ITS). Our results placed the five included species of Rhyncholacis , one of which is newly described herein, in a moderately well supported (73 BP) clade with M. clavigera . These results support the transfer of M. clavigera to Rhyncholacis ( R . clavigera ); the nomenclatural changes are made. In addition, a new species of Rhyncholacis ( R. paulana C.T. Philbrick & C.P. Bove) is illustrated and described. Rhyncholacis paulana is distinguished from all other species in the genus by its simple pinnately lobed leaf, which is fleshy and undulate. All other species of Rhyncholacis have leaves that are pinnately lobed, the lobes of which are repeatedly divided, or pinnately compound and characterized by finely dissected pinnate segments.
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A New Genus of Podostemaceae from Venezuela
Novon, 2011Co-Authors: C. Thomas Philbrick, Jacqueline Malecki, Nicholas P. Tippery, Hannah StevensAbstract:Abstract. The new genus Autana C. T. Philbrick and species A. andersonii C. T. Philbrick are illustrated and described. The new species is documented from four rivers that drain into the Orinoco River, Amazonas, Venezuela. Autana is distinguished by three characters that do not occur in other Neotropical Podostemaceae. The upper surface of the flattened stem has an anastomosing pattern that is derived from a continuation of leaf margins. When the capsule is mature, the pedicel apex (and receptacle) is swollen and hollow. After anthesis, locations where the deciduous stamens attached become darkened and shaped like an inverted teardrop. Phylogenetic analyses indicate A. andersonii is sister to a clade containing species of Castelnavia Tul. & Wedd., Oserya Tul. & Wedd., Noveloa C. T. Philbrick, and Rhyncholacis Tul., as well as upright stemmed species of Apinagia Tul., and Central American and Mexican species of Marathrum Bonpl.
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Monograph of Castelnavia (Podostemaceae)
Systematic Botany, 2009Co-Authors: C. Thomas Philbrick, Claudia Petean Bove, Thomas C. EdsonAbstract:Abstract A monograph of Castelnavia is presented. Phylogenetic analyses of morphological characters reveal eight nonhomoplasious synapomorphies supporting the monophyly of the genus: 1) absence of roots, a 2) unilocular mature, 3) anisolobous ovary that is 4) surrounded by stem tissue during and after anthesis, 5) horizontal at anthesis, with an 6) asymmetrically inflated pedicel apex and 7) longitudinal axis at 45–90° angle relative to pedicel axis, and 8) one deciduous capsule valve. Five species and two forms are recognized: Castelnavia fluitans, C. monandra, C. multipartita (C. m. forma multipartita, C. m. forma pendulosa C. T. Philbrick & C. P. Bove), C. noveloi, and C. princeps. Four species accepted by earlier authors are placed in synonymy. The morphology and ecology of the genus is discussed, and species descriptions, illustrations, a distribution map, lists of specimens examined, and a key to species are presented. Castelnavia occurs primarily in Brazil with one species in Bolivia. The greatest ...
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A New Species of Castelnavia (Podostemaceae) from Tocantins, Brazil
Novon, 2008Co-Authors: C. Thomas Philbrick, Claudia Petean BoveAbstract:Castelnavia noveloi C. T. Philbrick & C. P. Bove is distinguished from other species of Castelnavia Tulasne & Weddell (Podostemaceae) based on the production of stems arising perpendicular from the leaf petiole (petiolar stems) and the flattened rachis of the pinnate leaves. The new species is only known from rio Taquarussu, east of the town of Taquarussu, Tocantins, Brazil.
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Diamantina lombardii - an odd Brazilian member of the Podostemaceae
Flora, 2005Co-Authors: Rolf Rutishauser, R. Alejandro Novelo, Evelin Pfeifer, C. Thomas PhilbrickAbstract:Abstract This paper complements the diagnosis of the recently described genus Diamantina and its only species Diamantina lombardii Novelo, Philbrick and Irgang from Minas Gerais (Brazil). Four new features not known from other Podostemaceae–Podostemoideae are documented by microtome sections and SEM graphs: (i) The digitate foliage leaves lack vascular tissue completely. (ii) Leafy shoots produce one or two flowers in terminal and subterminal position. The spathella subtending the subterminal flower is scale-like and positionally homologous to a digitate bract (leaf), whereas the spathella covering the terminal flower bud is tubular (as usual for Podostemoideae). (iii) The usually rudimentary androecium consists of two stamens which form a complete whorl together with three inconspicuous tepals around the gynophore. (iv) The bilocular ovary has an apical cleft. Each carpel tip (hollow inside) is topped by a prominent horn-like stigma. Additional peculiar features of D. lombardii (already mentioned in Philbrick et al., 2004. Syst. Bot. 29, 109–117) are shown: presence of a prominent gynophore (mainly known from African Podostemoideae), and digitate leaves (as found in Cladopus from Eastern Asia to NE Australia).
R. Alejandro Novelo - One of the best experts on this subject based on the ideXlab platform.
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Diamantina lombardii - an odd Brazilian member of the Podostemaceae
Flora, 2005Co-Authors: Rolf Rutishauser, R. Alejandro Novelo, Evelin Pfeifer, C. Thomas PhilbrickAbstract:Abstract This paper complements the diagnosis of the recently described genus Diamantina and its only species Diamantina lombardii Novelo, Philbrick and Irgang from Minas Gerais (Brazil). Four new features not known from other Podostemaceae–Podostemoideae are documented by microtome sections and SEM graphs: (i) The digitate foliage leaves lack vascular tissue completely. (ii) Leafy shoots produce one or two flowers in terminal and subterminal position. The spathella subtending the subterminal flower is scale-like and positionally homologous to a digitate bract (leaf), whereas the spathella covering the terminal flower bud is tubular (as usual for Podostemoideae). (iii) The usually rudimentary androecium consists of two stamens which form a complete whorl together with three inconspicuous tepals around the gynophore. (iv) The bilocular ovary has an apical cleft. Each carpel tip (hollow inside) is topped by a prominent horn-like stigma. Additional peculiar features of D. lombardii (already mentioned in Philbrick et al., 2004. Syst. Bot. 29, 109–117) are shown: presence of a prominent gynophore (mainly known from African Podostemoideae), and digitate leaves (as found in Cladopus from Eastern Asia to NE Australia).
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DESARROLLO DE LOS VERTICILOS SEXUALES DE VANROYENELLA PLUMOSA NOVELO & Philbrick (PODOSTEMACEAE)
Acta Botanica Mexicana, 2001Co-Authors: Guillermina Murguia Sanchez, C. Thomas Philbrick, R. Alejandro Novelo, G. Judith Márquez GuzmánAbstract:The vegetative body of Vanroyenella plumosa Novelo & Philbrick (subfamily Podostemoideae) is composed of roots, leaves and a dorsiventrally flattened thalloid stem which, in the reproductive stage of the life cycle, develops cavities containing fascicles of flowers. In this work the embryology of this species was studied. Floral development depends on the flower position in the vegetative body of the plant, the plant position on the rock and the water level in the river. Vanroyenella plumosa possesses two stamens with tetrasporangiate anthers; the anther wall formation is of the basic type. Tetrads are tetrahedral, and the pollen is bicellular. The gynoecium is bicarpelar and bilocular with axial placentation and has two stigmas with idioblasts whose vacuoles contain proteins and carbohydrates. The ovules are anatropous, bitegmic and tenuinucellate. The embryo sac is monosporic in its origin and tetracellular of the Apinagia type. In Vanroyenella floral development occurs in the interior of the stem while the plant is submerged, and it is during this very early stage that the male and female gametes are formed. When the water level recedes, the pedicels elongate and anthesis occurs above water. The ovary wall supports the seed development after the rest of the vegetative tissue has died.
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Taxonomy of Mexican Podostemaceae
Aquatic Botany, 1997Co-Authors: R. Alejandro Novelo, C. Thomas PhilbrickAbstract:Abstract A taxonomic treatment of Podostemaceae in Mexico is presented, based on herbarium study and extensive field collections. Five genera and eight species of Podostemaceae (including Tristichaceae) occur in Mexico: Marathrum Humb. & Bonpl. (M. rubrum Novelo & Philbrick, M. schiedeanum (Cham.) Tul., M. tenue Liebm.), Oserya Tul. & Wedd. (O. coulteriana Tul., O. longifolia Novelo & Philbrick), Podostemum Michx. (P. ricciiforme (Liebm.) P. Royen), Tristicha Du Petit-Thouars (T. trifaria (Bory ex Willd.) Spreng.) and Vanroyenella Novelo and Philbrick (V. plumosa Novelo & Philbrick). More species occur on the Pacific slope (especially the states of Jalisco, Michoacan and Guerrero) than the Atlantic slope. In Mexico, Podostemaceae occur predominantly in three main climate types (A, hot and humid; B, dry; C, temperate and humid), and eight Mexican physiographic provinces. Podostemaceae are found primarily in river rapids that occur in the following vegetation types: oak-pine forest, moist mountain forest, tropical rain forest, tropical subdeciduous forest and tropical deciduous forest.
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Comparative pollen morphology of five New World genera of Podostemaceae
Aquatic Botany, 1997Co-Authors: Sean P. O'neill, C. Thomas Philbrick, Jeffrey M. Osborn, R. Alejandro NoveloAbstract:Abstract The Podostemaceae are the largest family of strictly aquatic angiosperms; however, relatively little is known about the palynology of the family. Pollen micromorphology and ultrastructure of five representative New World taxa are described, including Marathrum rubrum Novelo & Philbrick, Oserya coulteriana Tul., Podostemum ceratophyllum Michx., Tristicha trifaria (Bory ex Willd.) Sprengel, and Vanroyenella plumosa Novelo & Philbrick. Pollen grains from all five species are relatively small, spherical, microechinate, have a tectate-granular sexine and a thick nexine in non-apertural regions, and a semitectate sexine and a thin nexine in apertural regions. Characters that vary among the taxa include dispersal unit (monads or dyads), sculptural element morphology, infratectal granule size, and aperture morphology and ultrastructure. This is the first study to describe the pollen morphology of these five taxa in detail, and it is the first to illustrate the ultrastructure of pollen wall characters for any member of the family.
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A new species of Oserya (Podostemaceae) from Jalsico, Mexico
Novon, 1995Co-Authors: R. Alejandro Novelo, C. Thomas PhilbrickAbstract:A new species of Podostemaceae, Oserya longifolia, from Jalisco, Mexico, is described and illustrated. Leaf length is the most prominent feature that distinguishes this species. RESUMEN. Se describe e ilustra una especie nueva de la familia Podostemaceae, Oserya longifolia, del estado de Jalisco, M6xico. La longitud de la hoja es la caracteristica mas prominente que distingue a esta especie. Oserya Tulasne & Weddell is an American genus of six species, most recently monographed by van Royen (1954). Like all Podostemaceae, species of Oserya grow attached to rocks in swift-moving river currents. Five of the six species (O. biceps Tulasne & Weddell, O. flabelifera Tulasne & Weddell, O. minima van Royen, O. perpusilla (Went) van Royen, O. sphaerocarpa Tulasne) occur in northern Brazil and northeastern South America, while O. coulteriana Tulasne occurs in Mexico. Little is known of the geographic distributions of species of Oserya. In fact, four of the South American species are known only from the type collections. At the time of van Royen's monograph, O. coulteriana was known from few collections in the Mexican states of Jalisco and Michoacan. During our recent studies of Mexican Podostemaceae we have found this species to be relatively widespread in Colima, Nayarit, and Guerrero, in addition to Jalisco and Michoacin. A collection from southern Sinaloa is also now known (Novelo & Philbrick, unpublished). While conducting field studies of Mexican Podostemaceae, we made collections that did not correspond to any described species. These collections serve as the basis for the description of a new species, Oserya longifolia Novelo & Philbrick. serya longifolia Novelo & Philbrick, sp. nov. TYPE: Mexico. Estado de Jalisco: municipio de La Huerta, Rio Purificaci6n, 4 km al oeste de La Huerta, 19?30'N, 104?40'W, 300 m, 26 Mar. 1993, Novelo & Philbrick 1166 (holotype, MEXU; isotypes, MO, NY, WCSU). Figure 1. Herbae aquaticae plerumque caulibus stoloniferis applanatis. Folia circinata usque ad 40 cm longa; petiolus laevis; lamina repetite 2-4(-5)-divisa, divisionibus ultimis 0.3-0.6 mm latis, applanatis, apice acutis. Flores hermaphroditi, zygomorphi, pedicellati, axillares, solitarii. Tepala 3, filamentis alternata, uno in furca duorum staminum affixo. Stamina 2, andropodio portata; antherae ellipticae, dorsifixae, per 2 rimas laterales longitudinaliter dehiscentes. Ovarium 2 carpellis, 2 stylis 0.7-1.2 mm longi. Fructus 2 valvis, quaque valva 3-costata. Aquatic herbs usually with stoloniferous and flattened stems 0.8-1.0 mm diam., strongly adhering to rocks. Leaves alternate, circinate, up to 40 cm long, petiole 2-17 cm long, cylindrical, smooth, 0.7-1.6 mm diam., with a broadened base; blade 2-4(-5) divided, the ultimate divisions 0.3-0.6 mm wide, flattened, apex acute. Flowers hermaphroditic, zygomorphic, pedicellate, borne singly, axillary, protected by a spathella; spathella up to 7 mm long, thin clavate. Pedicels 3.5-8.0 mm long, elongating during anthesis, not expanded at capsule base. Tepals 3, alternate with the filaments, free, membranous, subulate, one of them attached in the fork between the two stamens; lateral tepals 0.9-1.2 mm long, medial tepal 0.7-0.9 mm long. Stamens 2, borne by an andropodium, andropodium 0.8-1.3 mm long, elongating during anthesis; filaments 1.01.8 mm long, subulate, flattened, elongating during anthesis, deciduous; anthers 0.4-0.6(-0.8) mm long, elliptic, dorsifixed, 2-celled, dehiscing longitudinally by 2 lateral slits. Ovary superior, 1.7-2.1 mm long, NovoN 5: 54-56. 1995. This content downloaded from 207.46.13.128 on Tue, 06 Sep 2016 05:39:13 UTC All use subject to http://about.jstor.org/terms Volume 5, Number 1 Novelo & Philbrick 55 1995 Oserya longifolia
Guillermina Murguia Sanchez - One of the best experts on this subject based on the ideXlab platform.
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desarrollo de los verticilos sexuales de vanroyenella plumosa novelo Philbrick podostemaceae
Acta Botanica Mexicana, 2001Co-Authors: Guillermina Murguia Sanchez, Alejandro R Novelo, Thomas C Philbrick, Judith Marquez G GuzmanAbstract:The vegetative body of Vanroyenella plumosa Novelo & Philbrick (subfamily Podostemoideae) is composed of roots, leaves and a dorsiventrally flattened thalloid stem which, in the reproductive stage of the life cycle, develops cavities containing fascicles of flowers. In this work the embryology of this species was studied. Floral development depends on the flower position in the vegetative body of the plant, the plant position on the rock and the water level in the river. Vanroyenella plumosa possesses two stamens with tetrasporangiate anthers; the anther wall formation is of the basic type. Tetrads are tetrahedral, and the pollen is bicellular. The gynoecium is bicarpelar and bilocular with axial placentation and has two stigmas with idioblasts whose vacuoles contain proteins and carbohydrates. The ovules are anatropous, bitegmic and tenuinucellate. The embryo sac is monosporic in its origin and tetracellular of the Apinagia type. In Vanroyenella floral development occurs in the interior of the stem while the plant is submerged, and it is during this very early stage that the male and female gametes are formed. When the water level recedes, the pedicels elongate and anthesis occurs above water. The ovary wall supports the seed development after the rest of the vegetative tissue has died.
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DESARROLLO DE LOS VERTICILOS SEXUALES DE VANROYENELLA PLUMOSA NOVELO & Philbrick (PODOSTEMACEAE)
Acta Botanica Mexicana, 2001Co-Authors: Guillermina Murguia Sanchez, C. Thomas Philbrick, R. Alejandro Novelo, G. Judith Márquez GuzmánAbstract:The vegetative body of Vanroyenella plumosa Novelo & Philbrick (subfamily Podostemoideae) is composed of roots, leaves and a dorsiventrally flattened thalloid stem which, in the reproductive stage of the life cycle, develops cavities containing fascicles of flowers. In this work the embryology of this species was studied. Floral development depends on the flower position in the vegetative body of the plant, the plant position on the rock and the water level in the river. Vanroyenella plumosa possesses two stamens with tetrasporangiate anthers; the anther wall formation is of the basic type. Tetrads are tetrahedral, and the pollen is bicellular. The gynoecium is bicarpelar and bilocular with axial placentation and has two stigmas with idioblasts whose vacuoles contain proteins and carbohydrates. The ovules are anatropous, bitegmic and tenuinucellate. The embryo sac is monosporic in its origin and tetracellular of the Apinagia type. In Vanroyenella floral development occurs in the interior of the stem while the plant is submerged, and it is during this very early stage that the male and female gametes are formed. When the water level recedes, the pedicels elongate and anthesis occurs above water. The ovary wall supports the seed development after the rest of the vegetative tissue has died.
James L Levenson - One of the best experts on this subject based on the ideXlab platform.
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Comprar Clinical Manual Of Psychosomatic Medicine | K. Philbrick | 9781585623938 | American Psychiatric Publishing
2020Co-Authors: K Philbrick, James R Rundell, P Netzel, James L LevensonAbstract:Tienda online donde Comprar Clinical Manual Of Psychosomatic Medicine al precio 53,56 € de K. Philbrick | J. Rundell | P. Netzel | J. Levenson, tienda de Libros de Medicina, Libros de Psiquiatria - Psiquiatria General
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comprar clinical manual of psychosomatic medicine k Philbrick 9781585623938 american psychiatric publishing
2011Co-Authors: K Philbrick, James R Rundell, P Netzel, James L LevensonAbstract:Tienda online donde Comprar Clinical Manual Of Psychosomatic Medicine al precio 53,56 € de K. Philbrick | J. Rundell | P. Netzel | J. Levenson, tienda de Libros de Medicina, Libros de Psiquiatria - Psiquiatria General
Judith Marquez G Guzman - One of the best experts on this subject based on the ideXlab platform.
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desarrollo de los verticilos sexuales de vanroyenella plumosa novelo Philbrick podostemaceae
Acta Botanica Mexicana, 2001Co-Authors: Guillermina Murguia Sanchez, Alejandro R Novelo, Thomas C Philbrick, Judith Marquez G GuzmanAbstract:The vegetative body of Vanroyenella plumosa Novelo & Philbrick (subfamily Podostemoideae) is composed of roots, leaves and a dorsiventrally flattened thalloid stem which, in the reproductive stage of the life cycle, develops cavities containing fascicles of flowers. In this work the embryology of this species was studied. Floral development depends on the flower position in the vegetative body of the plant, the plant position on the rock and the water level in the river. Vanroyenella plumosa possesses two stamens with tetrasporangiate anthers; the anther wall formation is of the basic type. Tetrads are tetrahedral, and the pollen is bicellular. The gynoecium is bicarpelar and bilocular with axial placentation and has two stigmas with idioblasts whose vacuoles contain proteins and carbohydrates. The ovules are anatropous, bitegmic and tenuinucellate. The embryo sac is monosporic in its origin and tetracellular of the Apinagia type. In Vanroyenella floral development occurs in the interior of the stem while the plant is submerged, and it is during this very early stage that the male and female gametes are formed. When the water level recedes, the pedicels elongate and anthesis occurs above water. The ovary wall supports the seed development after the rest of the vegetative tissue has died.
