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Tamer Keceli - One of the best experts on this subject based on the ideXlab platform.
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anomodon longifolius anomodontaceae Bryopsida new to the bryophyte flora of turkey
Turkish Journal of Botany, 2010Co-Authors: Muhammet Oren, Guray Uyar, Tamer KeceliAbstract:Anomodon longifolius (Schleich. ex Brid.) Hartm. is reported for the first time from Turkey, bringing the total number of Anomodon species known from Turkey to 6. This record extends its distribution range to the Western Black Sea region of Turkey. The nearest localities to Turkey fo r this species are in Bulgaria, Greece, the Caucasus, and Iran. This submeridional species was collected from 3 different localities (in Bolu and Bartin provinces) in Turkey. A site description, illustrations, and diagnostic characters of the Turkish specimens are given, together with notes on its distribution and ecology.
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bucklandiella microcarpa hedw bednarek ochyra ochyra grimmiaceae Bryopsida new to the moss flora of turkey and south west asia
Cryptogamie Bryologie, 2007Co-Authors: Gokhan Abay, Guray Uyar, Barbaros Cetin, Tamer KeceliAbstract:Bucklandiella microcarpa (Hedw.) Bednarek-Ochyra & Ochyra (Racomitrioideae, Grimmiaceae, Bryopsida) is recorded for the first time in Turkey. The specimens were collected from Kackar Mountains National Park in Rize province. The nearest localities of this species are in south Carpathians and Ural mountains. Therefore, this new record contributes to a remarkable range extension of this species towards south-west Asia.
Lars Hedenäs - One of the best experts on this subject based on the ideXlab platform.
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Scandinavian Oncophorus (Bryopsida, Oncophoraceae): species, cryptic species, and intraspecific variation
2017Co-Authors: Lars HedenäsAbstract:Lars Hedenäs (2017): Scandinavian Oncophorus (Bryopsida, Oncophoraceae): species, cryptic species, and intraspecific variation. European Journal of Taxonomy 315: 1-34, DOI: 10.5852/ejt.2017.31
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scandinavian oncophorus Bryopsida oncophoraceae species cryptic species and intraspecific variation
European journal of taxonomy, 2017Co-Authors: Lars HedenäsAbstract:Scandinavian members of the acrocarpous moss genus Oncophorus were revised after field observations had suggested unrecognized diversity. Based on molecular (nuclear: internal transcribed spacers 1 and 2, ITS; plastid: trn G UCC G2 intron, trn G, rps 4 gene + trn S- rps 4 spacer, rps 4) and morphological evidence, four morphologically distinguishable species are recognized, Oncophorus elongatus (I.Hagen) Hedenas, O. integerrimus Hedenas sp. nov. (syn. O. virens var. elongatus Limpr.), O. virens (Hedw.) Brid., and O. wahlenbergii Brid. ( O. sardous Herzog, syn. nov.). Oncophorus elongatus was earlier recognized, but much of its variation was hidden within O. wahlenbergii . Its circumscription is here expanded to include plants with long leaves having mostly denticulate or sharply denticulate upper margins and with long and narrow marginal cells in the basal portion of the sheathing leaf lamina. The new species O. integerrimus sp. nov. differs from O. virens in having more loosely incurved leaves and entire or almost entire upper leaf margins. Besides these characters, the species in the respective pairs differ in quantitative features of the leaf lamina cells. Several cryptic entities were found, in several cases as molecularly distinct as some of the morphologically recognizable species, and phylogeographic structure is present within O. elongatus and O. virens .
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phylogeny and morphological evolution of the amblystegiaceae Bryopsida
Molecular Phylogenetics and Evolution, 2002Co-Authors: Alain Vanderpoorten, Lars Hedenäs, Cymon J Cox, Jonathan A ShawAbstract:Abstract To circumscribe the moss family Amblystegiaceae, we performed a broad-scale analysis of trnL–trnF spacer sequence data for 168 species of the Hypnales and 11 species of the Hookeriales and additional analyses of trnL–trnF and atpB–rbcL (chloroplast DNA), one nuclear region, the internal transcribed spacers of 18S–26S rDNA, and 68 morphological characters for a reduced data set of 54 species of Hypnales. The traditionally circumscribed Amblystegiaceae are polyphyletic and include the Amblystegiaceae s. str. and the Calliergonaceae fam. nov., plus several taxa closely related to other Hypnalean families. Generic relationships within the redefined Amblystegiaceae were investigated by analyzing data from the three DNA regions and morphology as used in the broader analysis. Reconstruction of morphological evolution was evaluated using maximum-parsimony and maximum-likelihood. Numerous independent character-state transitions implied by the phylogeny suggest that morphological characters that have traditionally been used to delineate the Amblystegiaceae are homoplastic. Sporophytic traits, which are generally given primacy over gametophytic traits in moss classification, are more labile than previously thought, and many characters that are related to sporophyte specializations are strongly correlated with habitat conditions. The evolution of several gametophyte features previously thought to be reliable for delineating the family are also strongly correlated with habitat. These observations help to explain the instability of the Amblystegiaceae in previous taxonomic and phylogenetic analyses based on morphology.
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1 dichelyma capillaceum dicks myr Bryopsida fontinalaceae
Journal of Bryology, 1996Co-Authors: Lars Hedenäs, Susanne Godow, Kristoffer HylanderAbstract:Dioicous, male and female plants similar (Fig. 1). Plants medium-sized, irregularly pinnately or irregularly and sometimes sparsely branched, green or brownish green, not or very weakly glossy when dry. Stem leaves inserted in three distinct or sometimes indistinct rows, when moist slightly to moderately strongly falcato-secund, when dry slightly twisted, otherwise little changed, lanceolate, somewhat keeled (± V-shaped in transverse section), gradually narrowed to very long, ,narrowly acuminate apex, 3.26-6.12 mm long and 0.26-0.63 mm wide, upper 26-46% of leaf consisting entirely of the excurrent nerve, neither plicate nor transversely undulate; nerve single, longexcurrent, 42.0-77.5 Jlm wide near insertion, in transverse section plano-convex, 4-6stratose, cells ± homogeneous, surface cells on both adaxial and abaxial sides linear, similar to adjoining lamina cells, lateral sides of nerve near apex denticulate or usually coarsely denticulate or dentate; margin of lamina and lower part of excurrent nerve very weakly and obtusely denticulate to ± entire, plane, marginal 1-3(-4) cell rows of leaf lamina frequently bistratose, but otherwise not differentiated from other lamina cells; median lamina cells 48.3-252.0 x 5.0-8.8 Jlm, linear and slightly flexuose, with tapering or longly tapering ends, thin-walled or slightly incrassate, eporose, smooth; basal lamina cells shorter and slightly wider than mid-leaf cells, slightly to strongly incrassate, eporose or slightly porose; alar cells not different from other basal cells; in the ontogeny of the leaf lamina the basal cells are the last leaf cells to differentiate, and all basal cells differentiate ± simultaneously; shortly and narrowly decurrent; apical lamina cells linear, but usually somewhat shorter than mid-leaf cells, slightly incrassate, eporose. Branch leaves similar to stem leaves but slightly smaller, proximal branch leaves ovate and acute or shortly acuminate. Protonema often found on old leaves, especially on the nerve or sometimes on leaf margin, more rarely on other leaf portions, consisting of uniseriate, mostly unbranched threads of rectangular or shortly rectangular and slightly incrassate cells. Pseudoparaphyllia foliose, broadly ovate or broader than wide, with obtuse or rounded apex. Paraphyllia absent. Rhizoids inserted below leaf insertions, often on the branches, red-brown, not or hardly branched, smooth. Axillary hairs 2-3 in each axil, strictly axillary, with 2-4 upper cells, 7.8-10.5 Jlm wide, hyaline, and basal cells 1-2, rectangular to shortly linear, pale brown. Stem without central strand, cortex (including epidermis) of 3-4 layers of small and incrassate cells, basic tissue of large, thin-walled o~ slightly incrassate cells, no hyalodermis, flattened or slightly flattened. Perigonia lateral on stems and branches, perigonial leaves ovate, suddenly narrowed to obtuse,
Pôrto,kátia Cavalcanti - One of the best experts on this subject based on the ideXlab platform.
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Bryopsida from the cerrado and gallery forest of the Sete Cidades National Park, in the State of Piauí, Brazil
Sociedade Botânica do Brasil, 2002Co-Authors: Castro, Nívea Maria Carneiro Farias, Pôrto,kátia Cavalcanti, Yano Olga, Castro, Antônio Alberto Jorge FariasAbstract:Foi realizado o levantamento florístico das Bryopsida dos ecossistemas de cerrado e de mata ripícola do Parque Nacional de Sete Cidades localizado nos municípios de Piracuruca e Brasileira (04º0515'S e 41º3045'W), Piauí, Brasil. Foram identificadas 22 espécies de musgos pertencentes às famílias: Bartramiaceae (1 sp.), Bryaceae (2 sp.), Calymperaceae (2 sp.), Dicranaceae (3 sp.), Erpodiaceae (1 sp.), Fissidentaceae (6 sp.), Hypnaceae (1 esp.), Leucobryaceae (1 sp.), Pottiaceae (2 sp.), Sematophyllaceae (2 sp.) e Stereophyllaceae (1 sp.). Constituem-se novas referências para o Brasil, Weisiopsis nigeriana (Egun. & Olar.) Zand.; para o Nordeste, Campylopus heterostachys (Hampe) Jaeg. e para o Estado do Piauí: Bryum capillare Hedw., Bryum cruegeri Hampe ex C.Müller, Philonotis uncinata (Schwaegr.) Brid., Vesicularia vesicularis (Schwaegr.) Broth., Sematophyllum subsimplex (Hedw.) Mitt., Trichosteleum fluviale (Mitt.) Jaeg., Hyophila involuta (Hook.) Jaeg. & Sauerb., Calymperes palisotii Schwaegr. ssp. richardii (C.Müller) S.Edwards, Fissidens guianensis Mont., Fissidens intermedius C.Müller, Fissidens prionodesMont., Fissidens goyazensis Broth. eFissidens zollingeri Mont. Aos táxons são indicadas referências de descrições, ilustração e distribuição geográfica.This study is a floristic survey of the Bryopsida from the "cerrado" and from the gallery forest of the Sete Cidades National Park in the municipalities Piracuruca and Brasileira, in the State of Piauí, Brazil (04º0515'S and 41º30-45'W). Twenty-two taxa were determined from the following families: Bartramiaceae (1 esp.), Bryaceae (2 esp.), Calymperaceae (2 esp.), Dicranaceae (3 esp.), Erpodiaceae (1 esp.), Fissidentaceae (6 esp.), Hypnaceae (1 esp.), Leucobryaceae (1 esp.), Pottiaceae (2 esp.), Sematophyllaceae (2 esp.) and Stereophyllaceae (1 esp.). Weisiopsis nigeriana (Egun. & Olar.) Zand. is the first record of this species to Brazil, Campylopus heterostachys (Hampe) Jaeg. is the first of this species found in Brazilian northeast, and Bryum capillare Hewd., Bryum cruegeri Hampe ex C.Müller, Philonotis uncinata (Schwaegr.) Brid., Vesicularia vesicularis (Schwaegr.) Broth., Sematophyllum subsimplex (Hedw.) Mitt., Trichosteleum fluviale (Mitt.) Jaeg., Hyophila involuta (Hook.) Jaeg. & Sauerb., Calymperes palisotii Schwaegr. ssp. richardii (C.Müller) S.Edwards, Fissidens guianensis Mont., Fissidens intermedius C.Müller, Fissidens prionodes Mont., Fissidens goyazensis Broth. and Fissidens zollingeri Mont. are new to the State of Piauí. For each taxon a description is given along with references, illustrations and geographical distribution
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Levantamento florístico de Bryopsida de cerrado e mata ripícola do Parque Nacional de Sete Cidades, Piauí, Brasil
Sociedade Botânica do Brasil, 2002Co-Authors: Castro, Nívea Maria Carneiro Farias, Pôrto,kátia Cavalcanti, Yano Olga, Castro, Antônio Alberto Jorge FariasAbstract:Foi realizado o levantamento florístico das Bryopsida dos ecossistemas de cerrado e de mata ripícola do Parque Nacional de Sete Cidades localizado nos municípios de Piracuruca e Brasileira (04º0515'S e 41º3045'W), Piauí, Brasil. Foram identificadas 22 espécies de musgos pertencentes às famílias: Bartramiaceae (1 sp.), Bryaceae (2 sp.), Calymperaceae (2 sp.), Dicranaceae (3 sp.), Erpodiaceae (1 sp.), Fissidentaceae (6 sp.), Hypnaceae (1 esp.), Leucobryaceae (1 sp.), Pottiaceae (2 sp.), Sematophyllaceae (2 sp.) e Stereophyllaceae (1 sp.). Constituem-se novas referências para o Brasil, Weisiopsis nigeriana (Egun. & Olar.) Zand.; para o Nordeste, Campylopus heterostachys (Hampe) Jaeg. e para o Estado do Piauí: Bryum capillare Hedw., Bryum cruegeri Hampe ex C.Müller, Philonotis uncinata (Schwaegr.) Brid., Vesicularia vesicularis (Schwaegr.) Broth., Sematophyllum subsimplex (Hedw.) Mitt., Trichosteleum fluviale (Mitt.) Jaeg., Hyophila involuta (Hook.) Jaeg. & Sauerb., Calymperes palisotii Schwaegr. ssp. richardii (C.Müller) S.Edwards, Fissidens guianensis Mont., Fissidens intermedius C.Müller, Fissidens prionodesMont., Fissidens goyazensis Broth. eFissidens zollingeri Mont. Aos táxons são indicadas referências de descrições, ilustração e distribuição geográfica
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Briófitas de caatinga: 2. Agrestina, Pernambuco, Brasil
Sociedade Botânica do Brasil, 1996Co-Authors: Pôrto,kátia Cavalcanti, Bezerra,maria De Fátima De AndradeAbstract:Em uma área de caatinga do município de Agrestina (Lat. 8º27'28"S; Long. 35º57'13"W) - PE foram identificadas 13 espécies de Bryophyta, pertencentes as famílias Frullaniaceae, Lejeuneaceae, Ricciaceae - Hepaticopsida-Bryaceae, Dicranaceae, Ephemeraceae, Fissidentaceae, Leucobryaceae, Pottiaceae e Stereophyllaceae - Bryopsida. Constituem-se novos registros para a região Nordeste Cololejeunea minutíssima (Sm.) Schiffn. e Riccia brasiliensis Schiffn. e para Pernambuco Frullania dusenii Steph
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Floristic survey of epixylic Bryophytes of an area remnant of the Atlantic forest (Timbaúba - PE, Brazil) : 1., Hepaticopsida (except Lejeuneaceae) and Bryopsida
1996Co-Authors: Germano, Shirley Rangel, Pôrto,kátia CavalcantiAbstract:A survey of the epixylic bryoflora of a remaining Atlantic forest (seasonal coastal deciduous forest), Engenho Água Azul, in the municipality of Timbaúba ( Lat 7° 35S; Long. 35° 20W ), State of Pernambuco, Brazil, registered 35 species of bryophytes distributed in 11 families of Bryopsida: Calymperaceae, Pilotrichaceae, Fissidentaceae, Hookeriaceae, Hypnaceae, Leucobryaceae, Leucomiaceae, Orthotrichaceae, Plagiotheciaceae, Sematophyllaceae, Thuidiaceae, and 5 of Hepaticopsida: Aneuraceae, Frullaniaceae, Geocalycaceae, Plagiochilaceae and Radulaceae. New records northeastern of Brazil are: Frullania caroliniana Sullivant, F. gymnotis Nees & Mont., Ochrobryum stenophyllum Besch., Plagiochila trigonifolia Steph., Radula macrostachya Linbenb. & Gott., Riccardia digitiloba (Spruce ex Steph.) Hell, Thuidium tomentosum Schimp. ex Besch. and Trichosteleum pusillum (Hornsch.) Jaeg. There is an indication of the description, illustration, and geographic distribution for Brazil of all taxons found.En este trabajo se presenta el catálogo de la flora briofítica epífita de la selva tropical oceánica Engenho Agua Azul, situada en el municipio de Timbaúba (Lat. 7º 35S; Long. 35º 20O), perteneciente al estado de Pernambuco, Brasil. Se registraron 35 especies de briófitos distribuidas en 11 familias de la clase Bryopsida: Calymperaceae, Pilotrichaceae, Fissidentaceae, Hookeriaceae, Hypnaceae, Leucobryaceae, Leucomiaceae, Orthotrichaceae, Plagiotheciaceae, Sematophyllaceae, Thuidiaceae y 5 de la clase Hepaticopsida: Aneuracee, Frullaniaceae, Geocalycaceae, Plagiochilaceae y Radulaceae. Se aportan nuevas citas para la región nordeste del Brasil - Frullania caroliniana Sullivant, F. gymnotis Nees & Mont., Ochrobryum stenophyllumBesch., Plagiochila trigonifoliaSteph., Radula macrostachya Linbenb. & Gott., Riccardia digitiloba(Spruce ex Steph.) Hell, Thuidium tomentosum Schimp. ex Besch. yTrichosteleum pusillum (Horsnch.) Jaeg. Se presenta una descripción y una ilustración de cada taxon y se comenta su distribución geográfica
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Briófitas da caatinga 1: Estação Experimental do IPA, Caruaru - PE
Sociedade Botânica do Brasil, 1994Co-Authors: Pôrto,kátia Cavalcanti, Silveira,maria De Fátima Galdino Da, Sá,patrícia Sheyla De AlmeidaAbstract:Foi realizado o levantamento da brioflora de uma caatinga arbórea pertencente a Estação Experimental da Empresa Pernambucana de Pesquisas Agropecuárias (IPA), situada no município de Caruaru (8º08'00"-8º10'00"S, 36º02'00"-36º 10'00"W), Pernambuco. A área estudada compreende 20ha, tem altitude aproximada de 550m e médias anuais de precipitação e temperatura de 671 mm e 23ºC, respectivamente. As coletas foram realizadas durante as estações de estiagem (out-nov/1990) e chuvosa (mai-jun/1991), tendo sido investigados os seguintes substratos: solo, rocha, caules vivos e em decomposição. Registraram-se 16 espécies de briófitas, sendo 14 Bryopsida e 2 Hepaticopsida. Bryaceae e Fissidentaceae foram as famílias de maior representatividade específica na área. A maioria das coletas indicou a predominância de espécies terrícolas e saxícolas, seguindo-se as corticícolas e as epíxilas. O trabalho contém informações sobre distribuição geográfica e aspectos ecológicos das espécies
Castro, Antônio Alberto Jorge Farias - One of the best experts on this subject based on the ideXlab platform.
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Bryopsida from the cerrado and gallery forest of the Sete Cidades National Park, in the State of Piauí, Brazil
Sociedade Botânica do Brasil, 2002Co-Authors: Castro, Nívea Maria Carneiro Farias, Pôrto,kátia Cavalcanti, Yano Olga, Castro, Antônio Alberto Jorge FariasAbstract:Foi realizado o levantamento florístico das Bryopsida dos ecossistemas de cerrado e de mata ripícola do Parque Nacional de Sete Cidades localizado nos municípios de Piracuruca e Brasileira (04º0515'S e 41º3045'W), Piauí, Brasil. Foram identificadas 22 espécies de musgos pertencentes às famílias: Bartramiaceae (1 sp.), Bryaceae (2 sp.), Calymperaceae (2 sp.), Dicranaceae (3 sp.), Erpodiaceae (1 sp.), Fissidentaceae (6 sp.), Hypnaceae (1 esp.), Leucobryaceae (1 sp.), Pottiaceae (2 sp.), Sematophyllaceae (2 sp.) e Stereophyllaceae (1 sp.). Constituem-se novas referências para o Brasil, Weisiopsis nigeriana (Egun. & Olar.) Zand.; para o Nordeste, Campylopus heterostachys (Hampe) Jaeg. e para o Estado do Piauí: Bryum capillare Hedw., Bryum cruegeri Hampe ex C.Müller, Philonotis uncinata (Schwaegr.) Brid., Vesicularia vesicularis (Schwaegr.) Broth., Sematophyllum subsimplex (Hedw.) Mitt., Trichosteleum fluviale (Mitt.) Jaeg., Hyophila involuta (Hook.) Jaeg. & Sauerb., Calymperes palisotii Schwaegr. ssp. richardii (C.Müller) S.Edwards, Fissidens guianensis Mont., Fissidens intermedius C.Müller, Fissidens prionodesMont., Fissidens goyazensis Broth. eFissidens zollingeri Mont. Aos táxons são indicadas referências de descrições, ilustração e distribuição geográfica.This study is a floristic survey of the Bryopsida from the "cerrado" and from the gallery forest of the Sete Cidades National Park in the municipalities Piracuruca and Brasileira, in the State of Piauí, Brazil (04º0515'S and 41º30-45'W). Twenty-two taxa were determined from the following families: Bartramiaceae (1 esp.), Bryaceae (2 esp.), Calymperaceae (2 esp.), Dicranaceae (3 esp.), Erpodiaceae (1 esp.), Fissidentaceae (6 esp.), Hypnaceae (1 esp.), Leucobryaceae (1 esp.), Pottiaceae (2 esp.), Sematophyllaceae (2 esp.) and Stereophyllaceae (1 esp.). Weisiopsis nigeriana (Egun. & Olar.) Zand. is the first record of this species to Brazil, Campylopus heterostachys (Hampe) Jaeg. is the first of this species found in Brazilian northeast, and Bryum capillare Hewd., Bryum cruegeri Hampe ex C.Müller, Philonotis uncinata (Schwaegr.) Brid., Vesicularia vesicularis (Schwaegr.) Broth., Sematophyllum subsimplex (Hedw.) Mitt., Trichosteleum fluviale (Mitt.) Jaeg., Hyophila involuta (Hook.) Jaeg. & Sauerb., Calymperes palisotii Schwaegr. ssp. richardii (C.Müller) S.Edwards, Fissidens guianensis Mont., Fissidens intermedius C.Müller, Fissidens prionodes Mont., Fissidens goyazensis Broth. and Fissidens zollingeri Mont. are new to the State of Piauí. For each taxon a description is given along with references, illustrations and geographical distribution
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Levantamento florístico de Bryopsida de cerrado e mata ripícola do Parque Nacional de Sete Cidades, Piauí, Brasil
Sociedade Botânica do Brasil, 2002Co-Authors: Castro, Nívea Maria Carneiro Farias, Pôrto,kátia Cavalcanti, Yano Olga, Castro, Antônio Alberto Jorge FariasAbstract:Foi realizado o levantamento florístico das Bryopsida dos ecossistemas de cerrado e de mata ripícola do Parque Nacional de Sete Cidades localizado nos municípios de Piracuruca e Brasileira (04º0515'S e 41º3045'W), Piauí, Brasil. Foram identificadas 22 espécies de musgos pertencentes às famílias: Bartramiaceae (1 sp.), Bryaceae (2 sp.), Calymperaceae (2 sp.), Dicranaceae (3 sp.), Erpodiaceae (1 sp.), Fissidentaceae (6 sp.), Hypnaceae (1 esp.), Leucobryaceae (1 sp.), Pottiaceae (2 sp.), Sematophyllaceae (2 sp.) e Stereophyllaceae (1 sp.). Constituem-se novas referências para o Brasil, Weisiopsis nigeriana (Egun. & Olar.) Zand.; para o Nordeste, Campylopus heterostachys (Hampe) Jaeg. e para o Estado do Piauí: Bryum capillare Hedw., Bryum cruegeri Hampe ex C.Müller, Philonotis uncinata (Schwaegr.) Brid., Vesicularia vesicularis (Schwaegr.) Broth., Sematophyllum subsimplex (Hedw.) Mitt., Trichosteleum fluviale (Mitt.) Jaeg., Hyophila involuta (Hook.) Jaeg. & Sauerb., Calymperes palisotii Schwaegr. ssp. richardii (C.Müller) S.Edwards, Fissidens guianensis Mont., Fissidens intermedius C.Müller, Fissidens prionodesMont., Fissidens goyazensis Broth. eFissidens zollingeri Mont. Aos táxons são indicadas referências de descrições, ilustração e distribuição geográfica
Broutin Jean - One of the best experts on this subject based on the ideXlab platform.
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Presencia de Bryopsida fértil en los niveles Westfalianos del subgrup Itararé, Cuenca de Paraná, Brasil
2004Co-Authors: Amaral, Paula G. C., De Oliveira, Mary Bernardes, Ricardi-branco Fresia, Broutin JeanAbstract:The bryophyte fossils are rare, mainly in Paleozoic sedimentary rocks in spite of being present since the Silurian Period. In the Division Bryophyta, the fossils that belong to the Class Bryopsida are recognized since the Carboniferous, but they are extremely scarce. They are plentiful only in Permian sediments, in the Petchora, Kuznetsk and Russian Platform basins, also in Antarctica, Karoo basin (the last in South Africa) and India. Identified at the genus Dwykea, gametophyte specimens bearing pleurocarpous sporophyte were recovered from the lowermost levels of Itararé Subgroup, near Campinas city, S. Paulo State. These fossils correspond to the first register of bryophyte female gametophyte for the Carboniferous Period. The microflora in association with these fossils allow correlations of these levels to the Palynozone Ahrensisporites cristatus of Westphalian age. Related to proglacial sediments, they may correspond to a tundra vegetation covering the Northeastern border of Paraná Basin, during the Westphalian.Los fósiles atribuidos a briófitos son escasos, principalmente en rocas paleozoicas a pesar de ser registrados desde el Silúrico. Para la División Bryophyta, los fósiles correspondientes a la Clase Bryopsida comienzan a ser encontrados a partir del Carbonífero, aunque son bastante escasos. Registros más abundantes son conocidos para el Pérmico en rocas de las cuencas de Petchora, Kuznetsk y en la Plataforma Rusa, así como en la Antártica, en la Cuenca del Karoo (África del Sur) e India. Especímenes de gametófitos con esporofitos pleurocárpicos del género Dwykea fueron colectados en los niveles inferiores del Subgrupo Itararé, próximos a la ciudad de Campinas, Estado de S. Paulo. Estos fósiles corresponden al primer registro para el periodo Carbonífero de gametofitos femeninos fértiles. La microflora asociada a los fósiles de Dwykea permite establecer correlaciones con la Palinozona Ahrensisporites cristatus de edad westfaliana. Además los niveles donde fueron colectados los ejemplares de Dwykea corresponden a sedimentos proglaciales, que son interpretados como una vegetación de tundra que habitaba en el margen noreste de la Cuenca del Paraná, durante el Westfaliano
